I am not an expert, but from what I have read and seen with my car it seems that both fans run together. They can run at low speed or high speed, however, and they really go like crazy when they are on high speed. If they come on and your temperature is not over 100 degrees, then that is probably the low speed. If you ever get stuck in traffic on a hot day and your AC system is a bit low, so your fans don't come on at the low speed, then your temp may go up over 100 and then the fans will come on at high speed. I suppose they can also can come on at high speed without having low AC pressure, but I think it might take rather extreme conditions to get over 100 if your low speed fans are coming on with regularity due to the pressure in the AC system.
Others here are real experts, so they may chime in if I am getting too carried away with my newly acquired expertise on this matter.
Woops, I was too slow and a real expert answered while I was writing.
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I just couldn't give up on my 1995 E320.
I think it might be like always going back to that same bad relationship with an ex girlfriend.
You feel you love them too much, or you are just too stupid to know any better.
Flickr slideshow of my 1995 E320
http://www.flickr.com/photos/24145497@N06/sets/72157616572140057/